Kaessmann H. Origins, evolution, and phenotypic impact of new genes. Genome Res. 2010;20(10):1313-1326. doi:10.1101/gr.101386.109
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https://genome.cshlp.org/content/20/10/1313.full.pdf+html
Hypothesis of "out of testis"
Out of the testis: testis, compared with other organs, seem to play a more important role in permitting the nascent new genes to originate, and to subsequently adopt novel functions in other tissues by evolutionary time.
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